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The tree and shrub genus Zanthoxylum L., here regarded in its broad sense, including Fagara, is the largest of family Rutaceae, the citrus family. Approximately 200 species of Zanthoxylum are distributed mainly in tropical areas of the world, with few species reaching temperate latitudes. In the Neotropics, there are 77 species, 15 of them with subspecies, which adds a total of 105 taxa; its major center of species diversity is in the Andean-Amazonian region of Colombia, Ecuador, and Peru. The neotropical species of the genus are placed in four sections, Macqueria, Pterota, Tobinia, and Zanthoxylum, of which Macqueria contains the most morphologic variation and species, and is here divided into eight groups. This work, based on the examination of ca. 7500 herbarium specimens in ca. 45 world herbaria and field observation of 22 species, accounts for ca. 550 published names in Zanthoxylum and Fagara. All species are illustrated. It presents descriptions of all taxa, includes one new section of the genus, seven new species, three new subspecies, 22 new nomenclatural combinations, 91 new lectotypifications, two new neotypifications, a preliminary cladistic analysis based on morphology at the generic level, and an account of the wood anatomy of neotropical Zanthoxylum. Some of these plants have economic value as timber trees; medicinal compounds with promising value are also found throughout the genus.
Dr. Carlos Reynel was born March 15, 1957 in Lima, Peru, and grew up in the town of Chosica, in the western Andean piedmont, where he was educated in a state school. He attended the Facultad de Ciencias Forestales (FCF), Universidad Nacional Agraria-La Molina (UNALM) in Lima; in the 1980s, he worked as a field dendrologist, traveling and collecting trees in Amazonian and Andean Peru. He graduated as a forest engineer from UNALM in 1986, becoming a permanent professor there. In 1989 he traveled to the USA, where he completed his M.Sc. and Ph.D. studies in Biology at the University of Missouri-St. Louis, in its joint program with the Missouri Botanical Garden, under the supervision of Alwyn Gentry and P. Mick Richardson.
Dr. Reynel is currently staff professor at the FCF UNALM, where he is the director of the herbarium MOL woody collection, professor at UNALM Postgraduate School, member of the National Academy of Sciences of Peru, and research associate at the Missouri Botanical Garden. He has authored, with many collaborators, around fifteen books on miscellaneous aspects of woody plants of Peru, among them Árboles y arbustos andinos para agroforesterÍa y conservación de suelos (1990), El género Inga (Leguminosae) en el Perú (1997), Árboles útiles de la amazonÍa peruana (2003), Árboles útiles del ande peruano (2007), Árboles de los ecosistemas forestales andinos (2010), and Cómo se formó la diversidad ecológica del Perú (2013). Carlos Reynel's Author Page